Tel +44 (0) 1223 632850
Email subscriptions@biologists.com

Institutional subscriptions

Institutional prices

Tier coding grid

The Company of Biologists operates a tiered pricing system. If you are unclear about the tier in which your organisation should be classified, please contact subscriptions@biologists.com.

Tier 5 subscriptions and consortia agreements are negotiated individually with The Company of Biologists.

Subscriptions are accepted for a calendar year only.
All subscriptions are subject to the completion of a licence. By paying for your subscription, you are agreeing to our Copyright Policy and Terms and Conditions.


Personal subscriptions

Individuals and Societies

Individual/Society online subscription rates are only applicable when a subscription is for personal use.

  • Institutional, laboratory or shared use of any description is not permitted. Usage data is supplied to the publisher on a regular basis and abnormal usage is reported and will be investigated
  • By paying for your subscription, you are agreeing to our Copyright Policy and Terms and Conditions of use for personal subscribers

Subscriptions are accepted for a calendar year only.

  • VAT registered institutions and agents in the EU (excluding UK)
    VAT will not be charged on online and combined subscriptions, subject to the provision of a valid EU VAT number.
  • Non-VAT registered institutions, agents and individuals in the EU (excluding UK)
    VAT will be charged at the local rate on online and combined subscriptions.
  • UK institutions, agents and individuals
    VAT will be charged at 20% on all online and combined subscriptions.

The Company of Biologists Limited registered VAT number is GB 214 4240 10.

Full prepayment in the correct currency is required for all orders: USD ($) for orders delivered to USA or Canada; Euros (€) for orders delivered within Europe (excluding UK); GBP (£) for UK orders and orders delivered elsewhere.

The following methods of payment are accepted:

  • Our preferred payment method is via bank transfer to Barclays Bank PLC, Cambridge. Please see your invoice or contact subscriptions@biologists.com for the correct details.
  • By credit card we only accept MasterCard or Visa.
  • By cheque payable to ‘The Company of Biologists Limited’.

Online access is subject to the completion of a Site Licence.

All orders are regarded as firm and subscription payments are not refundable after the first issue has been dispatched or online access has been activated.

Institutional subscriptions

By purchasing a licence and by using the journals and websites of The Company of Biologists, you are agreeing to our Copyright Policy and the Terms and Conditions of use as set out in our Site Licence.

Site Licence

A single site is an organisational unit, which may be academic or non-academic and is defined to be within one metropolitan boundary, i.e. within a city.
For organisations located in more than one city, each city office is considered a different site and therefore a multi-site licence may be required.

Personal subscriptions

By purchasing a licence and by using the journals and websites of The Company of Biologists, you are agreeing to our Copyright Policy and the Terms and Conditions.

Our commitment to you:

Subscriptions are valid between 1 January and 31 December of any given year.

If you purchase an online subscription, a quality electronic version of your selected journal will be available before paper publication and distribution. We will try to keep our website available at all reasonable times, subject to necessary maintenance and unforeseen events.

The rights we will grant:

Our journals are protected by copyright and our copyright statements and disclaimers will apply. Please note that we do not accept responsibility for the work of our authors and also that, because our organisation is not-for-profit, we require to limit our liability to the amount of your subscription fee.

Users may:

  • Copy, download and store full text articles for their personal scholarly research and educational use
  • Use articles for teaching purposes and any other purposes permitted by our copyright statement. Each item must carry appropriate acknowledgement of the authors, journal, year, volume, issue, page citation and publisher

Users may not:

  • Remove or alter the authors’ names or the Publisher’s copyright notices or other means of identification or disclaimers as they appear on the websites or in the journal content
  • Systematically make print or electronic copies or multiple extracts or make multiple copies of any part of the content for any purpose other than that permitted by the copyright statement
  • Prepare derivative works or download, mount or distribute any part of the content on any electronic system or network, including without limitation the Internet and the World Wide Web, other than the Secure Network, except when permission has been obtained from The Company of Biologists
  • Alter, abridge or otherwise modify the content or any part of them for any purpose whatsoever
  • Users may not use open proxies. This is strictly prohibited
  • Share passwords or accounts with friends or colleagues

Users must seek and obtain permission from The Company of Biologists prior to all possible uses above. Contact: permissions@biologists.com.

Unauthorised use may lead to the termination of your subscription. A claim for compensation may also be raised.

Users accessing the journal sites through a mobile device will be detected and automatically redirected to the mobile-optimised web interface. Access control will be the same as on the main journal websites. An institution’s subscription will give users access to all content through the institution’s wifi connection, with the added ability for users to access content when off campus using a simple authentication process.

Supported operating systems include IOS (iPhone and iPod touch), Blackberry OS and Android smartphones.

In the first instance all new enquiries should be directed to subscriptions@biologists.com

If appropriate, we will put you in contact with our local representative.

The Company of Biologists have exclusive country representatives/agents for Brazil, China and Japan.
We also have outsourced sales representation in Bangladesh, Hong Kong, India, Indonesia, Laos, Malaysia, Philippines, Republic of Korea, Singapore, Sri Lanka, Taiwan, Turkey and Vietnam.

The Company of Biologists
Bidder Building
Station Road
CB24 9LF
United Kingdom

Tel +44 (0) 1223 632850

For USA, please write to:

Journal name
Air Business Ltd
c/o Worldnet Shipping Inc
156-15 146th Avenue, 2nd Floor
Jamaica, NY 11434

For UK and rest of world, email subscriptions@biologists.com.

Please quote your subscription number when requesting a change of address.

If you require a back issue quotation, please contact subscriptions@biologists.com.


Claims for missing print issues should be made immediately after receipt of a subsequent issue or within three months of the journal despatch date.

Claims for damage or shortage, should be reported within three days of receipt of the journal issue, otherwise a claim cannot be considered.

Please contact subscriptions@biologists.com, providing the following information:

  • Your subscriber number
  • Your full name and address
  • The journal name
  • Relevant volume number
  • Issue number

Development, Journal of Cell Science and Journal of Experimental Biology offer FREE online access to developing economies by participating in the following initiatives.


Over 6400 journals are now available free of charge to developing countries through the HINARI programme, organised by the World Health Organization and major publishers worldwide. This is one of the largest collections of biomedical and health literature, made available to health institutions in 108 countries, areas and territories and so contributing to improved world health.


Online Access to Research in the Environment (OARE) enables developing countries to have access to one of the world’s greatest collections of research in the Environmental Sciences. OARE is an international public-private consortium coordinated by the United Nations Environment Programme (UNEP), Yale University and leading scientific publishers.
Over 2990 peer reviewed titles are now available in more than 100 low income countries.


AGORA was set up by the Food and Agriculture Organization of the UN (FAO) together with major publishers. It provides 1900 journals to institutes in 107 counties in the fields of food, agriculture, environmental science and social sciences.


HighWire offers access to countries appearing in the World Bank’s list of ‘low income economies’ plus Angola, Armenia, Azerbaijan, Djibouti, Georgia, Indonesia and Turkmenistan. Users do not need to sign up for this service as HighWire’s software automatically detects the country they are connecting from and grants access accordingly.

Perpetual access

The Company of Biologists is a HighWire Press affiliated publisher and fully supports the LOCKSS Programme, a multi-institutional, multi-publisher initiative to create a permanent web publishing and access system.

LOCKSS replicates the traditional model of libraries keeping physical copies of books, journals, etc. in their collections, making it possible for libraries to keep copies of digital materials long-term. When libraries have custody of the content, they need not pay continued access fees for post-cancellation access.

LOCKSS provides libraries with the open source software and support to build their own collections and acquire a copy of the assets they pay for. LOCKSS preserves all formats and genres of web-published content. The intellectual content which includes the historical context (look and feel) is preserved. LOCKSS is OAIS compliant; the software migrates content forward in time; and the bits and bytes are continually audited and repaired.

Content preserved by libraries through LOCKSS becomes a part of their collection and they have perpetual access to 100% of titles preserved in their LOCKSS box.

Please see www.lockss.org for more information and to install a LOCKSS box.

Dark archive

The Company of Biologists has also partnered with CLOCKSS to preserve all their electronic journals at their geographically distributed network of redundant archive nodes located at 12 major research libraries around the world. This gives libraries the assurance that they will have a secure preservation solution and permanent access to our journals should a ‘trigger event’ ever occur.

Institutional subscriptions

All online subscriptions are subject to the completion of a site licence agreement. The link to our standard site licence is shown below:

Your online subscription will be fully activated following payment. Please ensure that you send a complete list of your registered IP addresses to subscriptions@biologists.com. Only registered IPs will be able to access the subscribed content.

Some institutions use proxy servers to network their environment. This means that all the IP addresses of the computers people are using appear to our server as a single IP address. Under the terms of our site licence, we permit the use of proxy servers on the condition that institutions deploy two factor authentication, specifically that each Authorised User may only access the Licensed Materials by inputting either: (i) a username and a password (ii) a staff ID and a PIN or (iii) an email address and a software token. Proxy servers that only require a staff ID to log in (without password) are not allowed for authenticating users.

Institutions within the same metropolitan area are classed as a single site and will be allocated to a Tier according to various factors listed in the Tier Coding Grid.
Institutions that operate from multiple national and international locations need to ensure that they have been accredited to the correct subscription tier with The Company of Biologists.

List each IP address, or partial IP address, following these rules:

  • The only characters allowed are numbers, dashes (-), or asterisks (*)
  • There should be 4 segments (containing max 3 characters) and no segment of an IP address can be blank
  • The allowed values for the first segment are 1-223
  • The allowed values for segments 2, 3, 4 are 0-254
  • The first segment cannot contain a range (-)
  • The first and second segments cannot contain a wildcard (*)
  • The third and fourth segments cannot both contain a range (-)

Example using both the wildcard character (*) and the range character (-):

Main site: All addresses beginning with 10.175.4 – use asterisk, in this case 10.175.4.*
Department: All addresses between 192.168.55.* and 192.168.60.* – use asterisk, in this case 192.168.55-60.*
Department: All addresses between and – use dashes, in this case

Update IP addresses
In the event that your institute’s IP addresses require amending or should you wish to expand your IP range over and above that currently registered, please contact subscriptions@biologists.com quoting your subscriber number.

Usage reports for the journals are COUNTER (www.projectcounter.org) compliant. Counting Online Usage of Networked Electronic Resources (COUNTER) is an international initiative serving librarians, publishers and intermediaries by setting standards that facilitate the recording and reporting of online usage statistics in a consistent, credible and compatible way.

The Company of Biologists currently provide COUNTER 4 reports. The provision of COUNTER 5 reports will start in February 2019. The new style of counting is very different to the previous method and therefore comparison year-on-year may be challenging.

Development usage reports
Journal of Cell Science usage reports
Journal of Experimental Biology usage reports

Shibboleth is a means of allowing access to subscribed content and is increasingly used by libraries signed up to their National Federation. The Company of Biologists has made Shibboleth available through HighWire for the UK, Germany, USA, Australia and Japan.

Users do not require individual usernames and passwords to gain access to journal material, as access is arranged by IP address.
Librarians or institutional administrators may change their usernames, passwords or contact details by following the links below:

How to change your password or contact details
Forgotten your username and/or password

Institutional usernames and passwords are for the use of the librarian or institutional administrator only and should not be shared with library users.

Personal subscriptions

Following payment of your online subscription in full, you will receive an email confirming your subscriber number. You will need this in order to activate your online subscription. You will only need to do this once.

You will create your own password during the activation process. You are the only one licenced to use this password.

You may reset your password by clicking the following link Forgotten user name/password

Institutional, laboratory or shared use of any description is not permitted. Usage data is supplied to the publisher on a regular basis and abnormal usage is reported and will be investigated.

Activate your subscription

By paying for your subscription, you are agreeing to our Copyright Policy  and Terms and Conditions of use for individual subscribers.


Institutional subscriptions

Please contact our customer services team at subscriptions@biologists.com providing the following information:

1. Journal name
2. Full name and address of organisation
3. Number of site locations
4. FTE
5. Tier Coding (if known)

When this happens, the IP address for your machine is not being recognised by our server. This could be because:

  • Your institutional subscription has expired or has not yet been activated
  • The person who activated the online subscription has not registered all the necessary IP addresses for your institution
  • The person who activated the online subscription does not realise that some subnets of your institution are routed through a proxy server that has not been authorised by The Company of Biologists

In the first instance talk to your librarian and let them know you are having difficulties or email  subscriptions@biologists.com.

If your institution has a subscription you will automatically have access to the Tables of Contents, Abstracts, Full Text searching, Full Text display, PDFs, Medline and GenBank links, and Tables of Contents in progress.

You will also see your institutional name or logo at the top of the page.

The subscription fee allows for unrestricted internet access at one location unless a multi-site licence has been established. Any user connecting from an authorised computer on your institutional network will be allowed access to the journals online. This includes members of the public accessing via terminals located onsite and under the control of the subscribing institution. Authorised users may also use home computers and other mobile devices provided such use can be authenticated by the institution. Institutional proxy servers must be authorised under the Terms and Conditions of the Site Licence.

A single site is an organisational unit, which may be academic or non-academic and is defined to be within one metropolitan boundary, i.e. within a city.
For organisations located in more than one city, each city office is considered a different site and therefore a multi-site licence may be required.

You will have access to all back content of the database; only the current unsubscribed period will be unavailable.

The licensed materials may not be used for any fee paid service, the systematic supply or distribution of the journals in any form or any similar commercial activity to anyone other than an Authorised User.

Subject to the terms of our Licence, the Licensee is permitted to supply to the user of another library in the same country as the licensee for non-commercial use (whether by post, fax or secure intermediate electronic transmission, using Ariel or its equivalent, whereby the electronic file is deleted immediately after printing), for the purposes of research or private study, a single copy of an electronic original of an individual article forming part of the licensed material, subject to the following limitations:

• Licensee may fulfil occasional requests only from another library;
• For requests for copying beyond the scope of the paragraph above, the requesting library must seek permission from The Company of Biologists and may make a request for permission directly to The Company of Biologists by sending an email to permissions@biologists.com.

If the licensee is located in the USA, the Licensee agrees to fulfil such requests in accordance with guidelines developed from time to time by the National Commission on New technological Uses of Copyright Works (CONTU guidelines).

Redistribution of the licensed materials to non-authorised users in electronic or digital form is not permitted unless the prior written consent of The Company of Biologists has been obtained.


The Company of Biologists allows librarians* to make multiple paper and electronic copies of single articles which are published by The Company of Biologists without charge, provided that the following conditions are met:

• Copying is done by a member of staff of the university
• Copying is for distribution without profit to student members of that institution for educational purposes only
• Copying is from the original article

The Company of Biologists allows readers* to copy, download and print out, for private study, single articles which are published by The Company of Biologists, without permission and without charge beyond the actual cost of copying, provided the copying is done by the person requiring its use or by an employee of the institution to which he/she belongs. Electronic copies must keep the article in its original form and any extracts of articles must be retyped verbatim. They may not be placed on the public internet or linked to or from an external or third party website. All articles and extracts of articles must include the identity of the author, the title of the work and the journal from which it is extracted.

Subject to the terms of the Licence, and subject to the paragraph below, the Licensee may incorporate links to Licensed Materials in Electronic Reserves for the use by Authorised Users in the course of instruction at the Licensee’s institution, but not for Commercial Use.

• In Electronic Reserves the use is conditional upon employing persistent links to the Licensed Materials so that usage is registered on The Company of Biologists’ online platform each time a student views the work on reserve. Each link shall carry appropriate acknowledgement of the source, listing title and author of the extract, title and author of the work, and the publisher as The Company of Biologists. Links shall be deleted by the Licensee when they are no longer used for such purpose.

• “Electronic Reserves” means a collection of links to Licensed Materials (e.g. journal articles) stored on the Secure Network by the Licensee for use by students in connection with specific courses of instruction offered by the Licensee to its students.

*Note that where The Company of Biologists reproduces any material with the permission of other publishers, you must seek permission from the copyright holder before using such material in accordance with this section.

Except as expressly provided for under this Copyright Policy, all copyright and other rights of The Company of Biologists in and to the Licensed Materials are reserved.

Please see librarian password header under the Manage subscriptions tab

Please email subscriptions@biologists.com with a complete list of IP addresses to be registered and annotate any proxy IPs.

Personal subscriptions

Please contact our customer services team at subscriptions@biologists.com providing the following information:

• Journal name
• Full name and permanent home address

When you purchase an online subscription to our journals, you will receive an email containing your subscriber number which will instruct you on how to activate your subscription. You will create your own password during the activation process.

Personal subscriptions are for individual use and passwords cannot be shared. Institutional, laboratory or shared use of any description is not permitted. Usage data is supplied to the publisher on a regular basis and abnormal usage is reported and will be investigated.

You may reset your password by following the link forgotten user name/password

You cannot gain full access to our journals without purchasing a subscription licence.

Individual articles maybe purchased on a pay-per-view basis via HighWire and The Copyright Clearance Center.

Transitional agreements

The Company of Biologists is embracing the move towards open access with a series of pilot transitional agreements with leading consortia and academic institutions. The Read and Publish multi-year agreements will allow researchers at participating institutions to publish an unlimited number of OA articles and access our three hybrid journals in return for a cost-efficient fixed fee with a low annual increase.

For further information please contact Richard Grove subscriptions@biologists.com

+44(0)1223 632 650

The Company of Biologists and Jisc are excited to announce a two-year pilot transitional open access (OA) agreement from January 2020.

The ‘Read and Publish’ deal will permit researchers at participating institutions unlimited access to all three subscription journals from The Company of Biologists, including the full archive dating back as far as 1853, and will allow the accepted articles of corresponding authors to be published open access by default and without limit.

Matthew Freeman, Chairman of The Company of Biologists and Head of the Dunn School of Pathology, University of Oxford said:

“We are thrilled to sign our first transitional agreement with Jisc. As a small not-for-profit publisher, the establishment of Read & Publish deals strengthens our support for the biological community and represents an important step in our open access journey.”

The Company of Biologists is a not-for-profit publishing company dedicated to supporting and inspiring the biological community. We do this by:

  • publishing five specialist peer-reviewed journals: Development, Journal of Cell Science, Journal of Experimental Biology, Disease Models & Mechanisms (open access) and Biology Open (open access)
  • funding meetings and Workshops to encourage cross-fertilisation of interdisciplinary ideas
  • providing grants, ranging from support for societies to Travelling Fellowships for graduate students and post-doctoral researchers.

Kathryn Spiller, licensing manager at Jisc said:

“We are delighted that The Company of Biologists has joined the small group of not-for-profit publishers who are pushing the open access agenda in offering an affordable ‘read and publish’ agreement to UK universities for 2020.”

About Jisc

Jisc’s vision is for the UK to be the most digitally advanced education and research nation in the world. At its heart is the super-fast national research and education network, Janet, with built-in cyber security protection.  Jisc also provides technology solutions for its members (colleges, universities and research centres) and customers (public sector bodies), helps members save time and money by negotiating sector-wide deals and provides advice and practical assistance on digital technology. Jisc is funded by the UK higher and further education and research funding bodies and member institutions.

We will announce participating institutions on our website and on Twitter as they are confirmed.

Visit our journal websites

Development Journal of Cell Science The Journal of Experimental Biology Disease Models & Mechanisms Biology Open

© 2019 The Company of Biologists Ltd | Registered Charity 277992
Registered in England and Wales | Company Limited by Guarantee No 514735
Registered office: Bidder Building, Station Road, Histon, Cambridge CB24 9LF, UK